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Richard Neher

Position: Research Group Leader
Unit: Neher
Phone: +49 7071 601 1345
Picture of Neher, Richard

Education and positions held

since 10/2010

Max-Planck Research Group Leader

08/2007-08/2010 Post-Doctoral Fellow at the Kavli Institute for Theoretical Physics, University of California, Santa Barbara, USA

02/2000-05/2007 Graduate Studies at the University of Munich, Germany


Undergraduate studies in physics at the University of Goettingen







Fellowships & Awards

12/2012 ARCHES award


ERC Starting Grant

08/2009 Harvey L. Karp Discovery Award

08/2007-08/2010 Post-Doctoral Fellow at the Kavli Institute for Theoretical Physics


Scholar of the Elite-Network of Bavaria







Professional Activities

since 08/2015 Associate Editor for Virus Evolution
since 08/2014 Member of the Board of Reviewing Editors of eLIFE

since 03/2011

Associate Editor for BMC Evolutionary Biology




Google Scholar profile


Peer-reviewed publications

  1. Predicting evolution from the shape of genealogical trees. Richard A Neher, Colin A Russel, Boris I Shraiman. eLIFE, in press
  2. Characterization of Genetic Diversity in the Nematode Pristionchus Pacificus from Population-Scale Resequencing Data. C Rödelsperger, RA. Neher, Andreas M. Weller, G Eberhardt, H Witte, WE Mayer, Dieterich, and RJ Sommer. Genetics. doi:10.1534/genetics.113.159855
  3. Genetic Diversity in the Interference Selection Limit. Benjamin H. Good, Aleksandra M. Walczak, Richard A. Neher, Michael M. Desai. PLoS Genetics, 10(3): e1004222.
  4. Quantifying the range of a lipid phosphate signal in vivo. Amrita Mukherjee, Richard A. Neher, Andrew D. Renault. J. Cell Science, epub
  5. Quantifying selection against synonymous mutations in HIV-1 env evolution. Fabio Zanini, Richard A. Neher. Journal of Virology, doi:10.1128/JVI.01529-13
  6. Coalescence and genetic diversity in sexual populations under selection Richard A. Neher, Taylor A. Kessinger, Boris I. Shraiman. PNAS 110 (39) 15836-15841
  7. Inferring HIV escape rates from multi-locus genotype data Taylor A. Kessinger, Alan S. Perelson and Richard A. Neher. Front. Immunol. 4:252. doi: 10.3389/fimmu.2013.00252
  8. Genetic draft, selective interference, and population genetics of rapid adaptation Richard A. Neher, Annual Reviews of Ecology, Evolution, and Systematics, vol 44.
  9. Genealogies of rapidly adapting populations Richard Neher, Oskar Hallatschek. PNAS 110 (2), 437-442
  10. Emergence of clones in sexual populations Richard Neher, Marjia Vucelja, Marc Mezard, Boris Shraiman. JSTAT 2013 (01), P01008
  11. Mathematical modeling of escape of HIV from cytotoxic T lymphocyte responses Vitaly Ganusov, Richard Neher, Alan Perelson. JSTAT 2013 (01), P01010
  12. Dynamic Mutation Selection Balance as an Evolutionary Attractor
    Sidhartha Goyal, Daniel J. Balick, Elizabeth R. Jerison, Richard A. Neher, Boris I. Shraiman, Michael M. Desai. Genetics (2012), Vol(191), pp. 1309-1319.
  13. FFPopSim: An efficient forward simulation package for the evolution of large populations
    Fabio Zanini and Richard A. Neher. Bioinformatics (2012), vol. 28 (24), pp 3332-3333
  14. Fluctuations of Fitness Distributions and the Rate of Muller's Ratchet
    R.A. Neher and B.I. Shraiman. Genetics (2012) Vol (191), pp. 1283-1293.
  15. Estimating the Strength of Selective Sweeps from Deep Population Diversity Data.
    P.W. Messer and R.A. Neher. Genetics (2012), Vol (191), pp. 593-605.
  16. Target search on a dynamic DNA molecule.
    T. Schoetz, R.A. Neher, and U. Gerland. Phys. Rev. E. (2011), Vol. 84, p. 051911.
  17. Statistical Genetics and Evolution of Quantitative Traits.
    R.A. Neher, B.I. Shraiman, Rev. Mod. Phys (2011), Vol 83(4), pp. 1283.
  18. Genetic Draft and Quasi-Neutrality in Large Facultatively Sexual Populations.
    R.A. Neher, B.I. Shraiman, Genetics (2011)
  19. Correlated Evolution of Nearby Residues in Drosophilid Proteins.
    B. Callahan, R.A. Neher, D. Bachtrog, P. Andolfatto, B.I. Shraiman, PLoS Genet (2011) vol. 7 (2) pp. e1001315
  20. Recombination rate and selection strength in HIV intra-patient evolution.
    R.A. Neher and T. Leitner. PLoS Comput Biol (2010) vol. 6 (1) pp. e1000660
  21. Rate of Adaptation in Large Sexual Populations.
    R.A. Neher, B. I. Shraiman and D. S. Fisher. Genetics (2010) vol. 184 pp. 467-481
  22. Competition between recombination and epistasis can cause a transition from allele to genotype selection
    R.A. Neher and B.I. Shraiman. Proc. Natl. Acad. Sci. USA, Vol. 106, pp. 6866-6871, 2009
  23. Blind source separation for the analysis of fluorescence microscopy images
    R.A. Neher, M. Mitkovski, E. Neher, F. Kirchhoff, F. Theis and A. Zeug. Biophys. J., Vol. 96, pp. 3791-3800, 2009.
  24. Blind Decomposition of Spectral Imaging Microscopy: A Study on Artificial and Real Test Data
    F. Theis , R. Neher, and A. Zeug. in ICA ’09: Proc. 8th Int. Conference on Independent Component Analysis and Signal Separation., pp. 548–556, 2009.
  25. Topological estimation of percolation thresholds
    R.A. Neher, K. Mecke, H. Wagner. JSTAT, P01011, 2008.
  26. Optimal stiffness for conformational transitions in macromolecules
    R.A. Neher, W. Moebius, E. Frey and U. Gerland. Phys. Rev. Lett., Vol. 99, p. 178101, 2007.
  27. Force-Induced DNA slippage
    F. Ku ̈hner, J. Morfill, R.A. Neher K. Blank and H. Gaub. Biophys. J., 92:2491-2497 (2007)
  28. Kinetic Accessibility of Buried DNA Sites in Nucleosomes
    W. Mo ̈bius, R.A. Neher and U. Gerland. Phys. Rev. Lett., Vol. 97, p. 208102, 2006.
  29. Intermediate phase in DNA melting
    R.A. Neher and U. Gerland. Phys. Rev. E., Vol. 73, p. R030902, 2006.
  30. DNA as a viscoelastic nanoelement
    R.A. Neher and U. Gerland. Biophys. J., Vol. 89, p. 3446-3855, 2005.
  31. Dynamics of Force-Induced DNA Slippage
    R.A. Neher and U. Gerland. Phys. Rev. Lett., Vol. 93, p. 198102, 2004.
  32. Applying Spectral Fingerprinting to the Analysis of FRET Images
    R.A. Neher and E. Neher, Micros. Res. Tech., Vol. 63, p.185-195, 2004.
  33. Optimizing imaging parameters for the separation of multiple labels in a fluorescence image
    R.A. Neher and E. Neher, J. of Microscopy, Vol. 213(1), p.46-62, 2004.